PARIS (AP) — With explosions and gunfire, security forces on Friday ended a three-day terror spree around Paris, killing the two al-Qaida-linked brothers who staged a murderous rampage at a satirical newspaper and an associate who seized a kosher supermarket to try to help them escape.

The worst terrorist attacks France has seen in decades left at least 20 people dead, including the three gunmen. The fate of a fourth suspect — a female accomplice of the market attacker — remained unclear.

At least seven people were killed Friday — the three terrorists and at least four hostages — a policewoman was killed Thursday and 12 people were massacred in the Paris newspaper attack Wednesday. Sixteen hostages were freed Friday, one from the printing plant where the two brothers were holed up and 15 others from the Paris grocery store.

Paris shut down a famed Jewish neighborhood amid fears that a wider terror cell might launch further attacks.

French President Francois Hollande urged his nation to remain united and alert. Calling the grocery attack "a terrifying anti-Semitic act," he said the terrorists were "fanatics who have nothing to do with the Islamic religion."

"The threats facing France are not finished," he said. "We must be vigilant."

The four attackers had ties to each other and to terrorism that reached back years and extended from Paris to al-Qaida in Yemen. They epitomized Western authorities' greatest fear: Islamic radicals who trained abroad and came home to stage attacks.

Said and Cherif Kouachi, the brothers who attacked the Charlie Hebdo satirical weekly, came out with guns blazing Friday evening after an all-day hostage siege at a printing plant northeast of Paris, a French police official said. They were killed and their hostage was freed, authorities said.

An accomplice, Amedi Coulibaly, took hostages Friday afternoon at a kosher grocery in the Porte de Vincennes neighborhood in Paris — then died in a nearly simultaneous raid there, said Gael Fabiano of the UNSA police union. Coulibaly had threatened to kill his hostages if French authorities launched an assault on the two brothers, a police official said.

A French television news network spoke directly with two of the terrorists Friday before they died. BFM television said it spoke with Cherif Kouachi as he and his brother were cornered near Charles de Gaulle airport and he told the station they were financed and dispatched by al-Qaida in Yemen.

The station also said it spoke with Coulibaly, who said the three men were coordinating and that he was with the militant Islamic State group. The organizations are normally rivals.

France has been on high alert since the Charlie Hebdo massacre left 12 people dead. The next day, a gunman shot a policewoman to death in a gunfight just south of Paris. Police later identified the gunman as Coulibaly, who had been a co-suspect with Cherif Kouachi in a court case involving terrorism that never made it to trial.

The Kouachi brothers led police on a chase around northeast France, robbing a gas station Thursday and stealing a car Friday morning before seizing hostages at a printing plant in Dammartin-en-Goele, a small industrial town near Charles de Gaulle airport.

Friday afternoon, explosions and gunshots rang out and white smoke rose outside as a police SWAT team clambered onto the roof. Audrey Taupenas, spokeswoman for the town near the Charles de Gaulle airport, said the brothers had died in the clash.

"They said they want to die as martyrs," Yves Albarello, a local lawmaker inside the command post, told French television station i-Tele.

Trying to fend off further attacks, the Paris mayor's office shut down all shops along Rosiers Street in the city's famed Marais neighborhood in the heart of the tourist district. Hours before the Jewish Sabbath, the street is usually crowded with shoppers — French Jews and tourists alike. The street is also only a kilometer (half a mile) from Charlie Hebdo's offices.

At the kosher grocery near the Porte de Vincennes neighborhood in Paris, the gunman burst in shooting just a few hours before the Jewish Sabbath began, declaring "You know who I am," the official recounted. The attack came before sundown when the store would have been crowded with shoppers.

Paris police released a photo of Coulibaly and his wife, Hayet Boumddiene, who the official said was his accomplice.

Several people wounded when the gunman opened fire in the grocery store were able to flee and get medical care. One hundred students were locked down in nearby schools and the highway ringing Paris was closed.

Cherif Kouachi, 32, was convicted of terrorism charges in 2008 for ties to a network sending jihadis to fight U.S. forces in Iraq.

A Yemeni security official said his 34-year-old brother, Said Kouachi, is suspected of having fought for al-Qaida in Yemen. Another senior security official said Said was in Yemen until 2012.

Both officials spoke to The Associated Press on condition of anonymity because of an ongoing investigation into Kouachi's stay in Yemen.

Both brothers were also on the U.S. no-fly list, a senior U.S. counterterrorism official said, speaking on condition of anonymity because the official was not authorized to discuss foreign intelligence publicly.


1:03 P.M. UPDATE

PARIS (AP) — Two al-Qaida-linked brothers suspected in the Charlie Hebdo massacre came out of their hideaway with guns blazing Friday and were killed in a clash with security forces, French police said. Moments later, another hostage-taker in Paris was killed in a separate clash, along with three of his hostages.

France has been high alert since the country's worst terror attack in decades — the massacre Wednesday in Paris at the satirical newspaper Charlie Hebdo that left 12 people dead.

After the two separate hostage-taking incidents began Friday, city officials scrambled to protect residents and tourists from further attacks, shutting down a famed Jewish neighborhood, putting schools under lockdown and urging residents to stay indoors and remain vigilant.

By Friday afternoon, explosions and gunshots rang out and white smoke rose up outside a printing plant in Dammartin-en-Goele, northeast of Paris, where brothers Cherif Kouachi, 32, and Said Kouachi, 34, had holed up with a hostage.

Security forces had surrounded the building for most of the day. After the explosions, a police SWAT forces were seen on the roof of the building and a police helicopter landed nearby it.

Audrey Taupenas, spokeswoman for the town near the Charles de Gaulle airport, said the brothers died in the clash. Their hostage was freed.

Another gunman who took at least five hostages Friday afternoon at a kosher grocery in Paris also died in a nearly simultaneous raid there, said Gael Fabiano of the UNSA police union.

The gunman, identified as Amedy Coulibaly, had earlier threatened to kill his hostages if French authorities launched an assault on the two brothers, a police official said. The two sets of hostage-takers know each other, said the official, who was not authorized to discuss the rapidly developing situation.

One other police officer said three hostages also died at the grocery. None could say what happened to the woman listed on a police bulletin as his accomplice.

Security forces stormed the Paris grocery near the Porte de Vincennes neighborhood minutes after news of clashes at the printing plant. Moments later, several people were seen being led out of the store. It was not clear exactly how many hostages had been inside or how many were freed.

Trying to fend off further attacks, the Paris mayor's office shut down all shops along Rosiers Street in the city's famed Marais neighborhood in the heart of the tourist district. Hours before the Jewish Sabbath, the street is usually crowded with shoppers — French Jews and tourists alike. The street is also only a kilometer (a half mile) away from Charlie Hebdo's offices.

The gunman had burst shooting into the kosher store just a few hours before the Jewish Sabbath began, declaring "You know who I am," the official recounted. The attack came before sundown when the store was crowded with shoppers. Several wounded people were able to flee, police said.

Paris police released photos of Coulibaly, also believed responsible for the roadside killing of a Paris policewoman, and one of a second suspect, a woman named Hayet Boumddiene, who they said was his accomplice.

About 100 students were placed under lockdown in schools nearby and the highway ringing Paris was closed.

Hours before and 40 kilometers (25 miles) away, a convoy of police trucks, helicopters and ambulances streamed toward Dammartin-en-Goele, a small industrial town near Charles de Gaulle airport, to seize the Charlie Hebdo suspects, who had hijacked a car in a nearby town after more than two days on the run.

"They said they want to die as martyrs," Yves Albarello, a local lawmaker inside the command post, told French television station i-Tele.

With the brothers trapped, Charles de Gaulle closed two runways to arrivals to avoid interfering in the standoff or endangering planes.

Both brothers had ties to terrorist networks. Cherif Kouachi was convicted of terrorism charges in 2008 for ties to a network sending jihadis to fight U.S. forces in Iraq.

A Yemeni security official said his brother, Said, is suspected of having fought for al-Qaida in Yemen. Another senior security official said Said was in Yemen until 2012. Both officials spoke to The Associated Press on condition of anonymity because of an ongoing investigation into Kouachi's stay in Yemen.

Both brothers were also on the U.S. no-fly list, a senior U.S. counterterrorism official said.

Nine people, members of the brothers' entourage, have been detained for questioning in several regions.

The satirical weekly Charlie Hebdo had long drawn threats for its depictions of Islam, although it also satirized other religions and political figures. The weekly paper had caricatured the Prophet Muhammad, and a sketch of Islamic State's leader was the last tweet sent out by the irreverent newspaper, minutes before the attack.

Eight journalists, two police officers, a maintenance worker and a visitor were killed in the newspaper attack, including the paper's editor. Charlie Hebdo plans a special edition next week, produced in the offices of another paper.

Authorities around Europe have warned of the threat posed by the return of Western jihadis trained in warfare. France counts at least 1,200 citizens in the war zone in Syria — headed there, returned or dead. Both the Islamic State group and al-Qaida have threatened France, home to Western Europe's largest Muslim population.


11:21 A.M. ORIGINAL STORY

PARIS (AP) — A security official says the two brothers suspected in the Charlie Hebdo massacre came out firing, prompting the assault on the building where they had holed up with a hostage.

The official was not authorized to speak about the sensitive operations and spoke on condition of anonymity.

Officials say the brothers died in the assault.

Another official, police union representative Christophe Crepin, said it appeared that the gunman who took hostages at a kosher market had also died in a nearly simultaneous raid there.

Crepin spoke to LCI television.

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